An analysis of concurrent sound-scattering records and biological collections in the sea show a correlation of the deeper and more intense scattering of sound with fishes and of the upper and lighter scattering with a dominant population of euphausiids. The main group of fishes concerned are the Myctophidae, which possess all of the essential qualifications of scatterers.
Tucker, Gordon H.. 1951. "Relation of fishes and other organisms to the scattering of underwater sound." Journal of Marine Research 10, (2). https://elischolar.library.yale.edu/journal_of_marine_research/749